22:speciation&18:speciation&12:speciation&8:speciation&6:speciation&4:speciation&0:leaf&3:creation&1:leaf&2:leaf&5:leaf&7:leaf&11:creation&9:leaf&10:leaf&17:speciation&15:duplication&13:leaf&14:leaf&16:leaf&21:speciation&19:leaf&20:leaf